Soldier Beetles: Difference between revisions
No edit summary |
m The LinkTitles extension automatically added links to existing pages (https://github.com/bovender/LinkTitles). |
||
(27 intermediate revisions by one other user not shown) | |||
Line 1: | Line 1: | ||
== Description & Overview == | == Description & Overview == | ||
<center> | <center> | ||
{| class="wikitable" style="margin-left:auto; margin-right:auto; background-color:#e9efda; | {| class="wikitable" style="margin-left:auto; margin-right:auto; background-color:#e9efda; | ||
|+ !colspan="7" style="min-width:12em; text-align: center; background-color: rgb(235,235,210)|Soldier Beetle Taxonomy <ref name= "bugguide">BugGuide. (2025). Family Cantharidae - Soldier Beetles. BugGuide. https://bugguide.net/node/view/118/tree.</ref> | |+ !colspan="7" style="min-width:12em; text-align: center; background-color: rgb(235,235,210)|Soldier Beetle Taxonomy <ref name= "bugguide">BugGuide. (2025). Family Cantharidae - Soldier Beetles. BugGuide. https://bugguide.net/node/view/118/tree.</ref> | ||
!colspan="7"|[[File:beetle2.jpg|501px|thumb|center|Soldier Beetle on a Plant in Chile (Photographed by Claudio Maureira, 2019) <ref name= "pics">iNaturalist. https://www.inaturalist.org/.</ref>.]] | !colspan="7"|[[File:beetle2.jpg|501px|thumb|center|Soldier Beetle (''Dysmorphocerus dilaticornis'') on a Plant in Chile (Photographed by Claudio Maureira, 2019) <ref name= "pics">iNaturalist. https://www.inaturalist.org/.</ref>.]] | ||
|- | |- | ||
! Kingdom !! Phylum !! Subphylum!! Class !! Order !! Suborder !! Family | ! Kingdom !! Phylum !! Subphylum !! Class !! Order !! Suborder !! Family | ||
|- | |- | ||
| Animalia || Arthropoda || Hexapoda || Insecta || Coleoptera || Polyphaga || Cantharidae | | Animalia || [[Arthropoda]] || Hexapoda || Insecta || Coleoptera || Polyphaga || Cantharidae | ||
|} | |} | ||
</center> | </center> | ||
Soldier beetles (Cantharidae) are the second largest group in the beetle (Coleoptera) superfamily Elateroidea, containing approximatley 5,500 species. These beetles are common throughout the temperate region of the northern hemisphere <ref name= "motyka">Motyka, M., Kusy, D., Biffi, G., Geiser, M., Kazantsev, S.V., Bilkova, R., Jahodarova, E., Vogler, A.P., Bocak, L.. (2023). Untangling the evolution of soldier beetles ([[Coleoptera]]: Cantharidae) and the evaluation of the morphological phylogenetic signal in a soft-bodied elateroid lineage. Cladistics. 39: 548-570. https://doi.org/10.1111/cla.12555.</ref>. | |||
Soldier beetle larvae tend to be darkly colored and look almost reptilian. As adults, soldier beetles grow to about half of an inch long and are yellow, orange, brown, or black in color <ref name= "hahn">Hahn, Jeffrey. (2023). Soldier beetles. University of Minnesota Extension. https://extension.umn.edu/yard-and-garden-insects/soldier-beetles#:~:text=Soldier%20beetles%20are%20beneficial%20insects&text=Because%20of%20their%20frequent%20contact,go%20away%20on%20their%20own.</ref>. | |||
Though soldier beetles are sometimes considered pests, they are beneficial, as they are a pollinator species <ref name= "hahn">Hahn, Jeffrey. (2023). Soldier beetles. University of Minnesota Extension. https://extension.umn.edu/yard-and-garden-insects/soldier-beetles#:~:text=Soldier%20beetles%20are%20beneficial%20insects&text=Because%20of%20their%20frequent%20contact,go%20away%20on%20their%20own.</ref>. Soldier beetles may be confused with certain species of blister beetles, fireflies, or glowworm beetles <ref name="ucipm">UCIPM. (2025). Soldier Beetles. UCIPM. https://ipm.ucanr.edu/natural-enemies/soldier-beetles/#gsc.tab=0.</ref>. | |||
== Ecology & Evolution == | == Ecology & Evolution == | ||
From midsummer to early fall, soldier beetles tend to pollinate yellow flowers of the family Asteraceae. Some adult soldier beetles feed on nectar | Soldier beetles have four life stages: Egg, Larva, Pupa, and Adult. Eggs are laid from late August to early September. Larvae are assumed to predate on small [[invertebrates]] within the [[soil]]. After winter, larvae are found moving through leaf litter, plant debris, and other areas of high humidity. Pupae form in early summer, and adults emerge starting in late July <ref name= "hahn">Hahn, Jeffrey. (2023). Soldier beetles. University of Minnesota Extension. https://extension.umn.edu/yard-and-garden-insects/soldier-beetles#:~:text=Soldier%20beetles%20are%20beneficial%20insects&text=Because%20of%20their%20frequent%20contact,go%20away%20on%20their%20own.</ref>. One notable adaption of soldier beetles is that select ones can produce compounds to help stave off predators, like spiders <ref name= "colorado">Colorado State University. (2017). Soldier Beetle. Western Colorado [[Insects]]. https://wci.extension.colostate.edu/wp-content/uploads/sites/14/2017/03/SoldierBeetles.pdf.</ref>. | ||
From midsummer to early fall, soldier beetles tend to pollinate yellow flowers of the family Asteraceae (such as dandelions and goldenrods). Some adult soldier beetles feed on nectar and pollen <ref name= "colorado">Colorado State University. (2017). Soldier Beetle. Western Colorado [[Insects]]. https://wci.extension.colostate.edu/wp-content/uploads/sites/14/2017/03/SoldierBeetles.pdf.</ref>. Other adults may prey on aphids and soft-bodied insects <ref name="ucipm">UCIPM. (2025). Soldier Beetles. UCIPM. https://ipm.ucanr.edu/natural-enemies/soldier-beetles/#gsc.tab=0.</ref>. | |||
Along with their important ecological role, soldier beetles have been significant in evolutionary studies. Currently, 25 species in 16 genera of soldier beetles have been discovered in fossiliferous amber resin. The oldest soldier beetle fossil discovered comes from Early Cretaceous Lebanese amber <ref name="esa">Entomology Today. New Soldier Beetle Species Found in 99-million-year-old Amber. Entomological Society of America. https://entomologytoday.org/2016/10/14/new-soldier-beetle-species-found-in-99-million-year-old-amber/.</ref>. | |||
Based on molecular dating, the Cantharidae family is thought to have originated 156.3 million years ago <ref name= "motyka">Motyka, M., Kusy, D., Biffi, G., Geiser, M., Kazantsev, S.V., Bilkova, R., Jahodarova, E., Vogler, A.P., Bocak, L.. (2023). Untangling the evolution of soldier beetles ([[Coleoptera]]: Cantharidae) and the evaluation of the morphological phylogenetic signal in a soft-bodied elateroid lineage. Cladistics. 39: 548-570. https://doi.org/10.1111/cla.12555.</ref>. | Based on molecular dating, the Cantharidae family is thought to have originated 156.3 million years ago <ref name= "motyka">Motyka, M., Kusy, D., Biffi, G., Geiser, M., Kazantsev, S.V., Bilkova, R., Jahodarova, E., Vogler, A.P., Bocak, L.. (2023). Untangling the evolution of soldier beetles ([[Coleoptera]]: Cantharidae) and the evaluation of the morphological phylogenetic signal in a soft-bodied elateroid lineage. Cladistics. 39: 548-570. https://doi.org/10.1111/cla.12555.</ref>. | ||
Line 30: | Line 32: | ||
===Cantharis=== | ===Cantharis=== | ||
[[File:beetle3.jpg|501px|thumb|center|Rustic Soldier Beetle (''Cantharis rustica'') On a Plant in France (Photographed by dromensis, 2025)]] | [[File:beetle3.jpg|501px|thumb|center|Rustic Soldier Beetle (''Cantharis rustica'') On a Plant in France (Photographed by dromensis, 2025) <ref name= "pics">iNaturalist. https://www.inaturalist.org/.</ref>.]] | ||
''Cantharis'' species are usually | ''Cantharis'' species are usually grayish-brown with a red or orange head and prothorax <ref name= "colorado">Colorado State University. (2017). Soldier Beetle. Western Colorado [[Insects]]. https://wci.extension.colostate.edu/wp-content/uploads/sites/14/2017/03/SoldierBeetles.pdf.</ref>. Beetles in this genus are commonly found in grasslands and forb fields <ref name="pelletier">Pelletier G. & Hebert, C. (2014). The Cantharidae of Eastern Canada and Northeastern United States. Canadian Journal of [[Arthropod]] Identification. https://doi.org/10.3752/cjai.2014.25.</ref>. ''Cantharis'' species are some of the largest soldier beetles, and adults can be between 9 to 13 millimeters long. These beetles have a red pronotum (back of head) with a black marking in the center. The top of these beetles' heads, as well as their wing covers, are black <ref name= "wildlife">The Wildlife Trusts. (2025). Soldier beetle. The Wildlife Trusts. https://www.wildlifetrusts.org/wildlife-explorer/invertebrates/beetles/soldier-beetle.</ref>. | ||
===Chauliognathus=== | ===Chauliognathus=== | ||
[[File:beetle4.jpg|501px|thumb|center|Plague Soldier Beetle (''Chauliognathus lugubris'') On a Plant in Australia (Photographed by zichen1, 2025)]] | [[File:beetle4.jpg|501px|thumb|center|Plague Soldier Beetle (''Chauliognathus lugubris'') On a Plant in Australia (Photographed by zichen1, 2025) <ref name= "pics">iNaturalist. https://www.inaturalist.org/.</ref>.]] | ||
Most ''Chauliognathus'' species are found in the western United States, Australia, and South America, excepting two species in the state of Virginia. Adults in this genus are | Most ''Chauliognathus'' species are found in the western United States, Australia, and South America, excepting two species in the state of Virginia. Adults in this genus are also some of the largest soldier beetles at 8 to 15 millimeters long. They are yellow to orange, with black coloration on their forewings and prothorax <ref name="catron">Catron, K.A.. (2021). Biology of Chauliognathus spp. (Hentz) (Coleoptera: Cantharidae) in Virginia agroecosystems. Virginia Polytechnic Institute and State University. https://vtechworks.lib.vt.edu/server/api/core/bitstreams/f6f5fd1b-f813-4947-a14a-1b06d989fc18/content.</ref>. Like ''Cantharis'', ''Chauliognathus'' species are also commonly found in grasslands and forb fields <ref name="pelletier">Pelletier G. & Hebert, C. (2014). The Cantharidae of Eastern Canada and Northeastern United States. Canadian Journal of [[Arthropod]] Identification. https://doi.org/10.3752/cjai.2014.25.</ref>. | ||
===Podabrus=== | ===Podabrus=== | ||
[[File:beetle6.jpeg|501px|thumb|center|Wrinkled Soldier Beetle (''Podabrus rugosulus'') On a Plant in South Carolina, USA (Photographed by xx7trey, 2024)]] | [[File:beetle6.jpeg|501px|thumb|center|Wrinkled Soldier Beetle (''Podabrus rugosulus'') On a Plant in South Carolina, USA (Photographed by xx7trey, 2024) <ref name= "pics">iNaturalist. https://www.inaturalist.org/.</ref>.]] | ||
Species in the ''Podabrus'' genus tend to feed on aphids and soft-bodied insects <ref name="podabrus">Montana State University. (2022). Podabrus sp. Montana State University. https://www.montana.edu/yellowstoneinsects/coleoptera/cantharidae/podabrus_sp.html#:~:text=Podabrus%20is%20a%20genus%20in,exclusively%20near%20bodies%20of%20water.</ref>. | Species in the ''Podabrus'' genus tend to feed on aphids and soft-bodied insects <ref name="podabrus">Montana State University. (2022). Podabrus sp. Montana State University. https://www.montana.edu/yellowstoneinsects/coleoptera/cantharidae/podabrus_sp.html#:~:text=Podabrus%20is%20a%20genus%20in,exclusively%20near%20bodies%20of%20water.</ref>. The larve of ''Podabrus'' genus are thought to reside in organic debris and the upper layers of soil <ref name="crowley">Crowley, L., Chua, P., & Kusy, D.. (2023). The genome sequence of a soldier beetle, Podabrus alpinus (Paykull, 1798). PubMed Central. https://doi.org/10.12688/wellcomeopenres.18890.1.</ref>. ''Podabrus'' adults typically range from 7 to 9 millimeters long. They tend to have leathery, flexible wing covers and are usually brownish-black with yellow to red sides and yellow faces <ref name="minnesota">MinnesotaSeasons.com. (2025). Wrinkled Soldier Beetle. MinnesotaSeasons.com. http://www.minnesotaseasons.com/Insects/wrinkled_soldier_beetle.html#:~:text=Podabrus%20rugosulus%20is%20a%20small,and%20rounded%20at%20the%20tip.</ref>. | ||
== References == | == References == | ||
<ref name= "motyka">Motyka, M., Kusy, D., Biffi, G., Geiser, M., Kazantsev, S.V., Bilkova, R., Jahodarova, E., Vogler, A.P., Bocak, L.. (2023). Untangling the evolution of soldier beetles ([[Coleoptera]]: Cantharidae) and the evaluation of the morphological phylogenetic signal in a soft-bodied elateroid lineage. Cladistics. 39: 548-570. https://doi.org/10.1111/cla.12555.</ref> | <ref name= "motyka">Motyka, M., Kusy, D., Biffi, G., Geiser, M., Kazantsev, S.V., Bilkova, R., Jahodarova, E., Vogler, A.P., Bocak, L.. (2023). Untangling the evolution of soldier beetles ([[Coleoptera]]: Cantharidae) and the evaluation of the morphological phylogenetic signal in a soft-bodied elateroid lineage. Cladistics. 39: 548-570. https://doi.org/10.1111/cla.12555.</ref> | ||
<ref name= "hahn">Hahn, Jeffrey. (2023). Soldier beetles. University of Minnesota Extension. https://extension.umn.edu/yard-and-garden-insects/soldier-beetles#:~:text=Soldier%20beetles%20are%20beneficial%20insects&text=Because%20of%20their%20frequent%20contact,go%20away%20on%20their%20own.</ref> | <ref name= "hahn">Hahn, Jeffrey. (2023). Soldier beetles. University of Minnesota Extension. https://extension.umn.edu/yard-and-garden-insects/soldier-beetles#:~:text=Soldier%20beetles%20are%20beneficial%20insects&text=Because%20of%20their%20frequent%20contact,go%20away%20on%20their%20own.</ref> | ||
<ref name= "colorado">Colorado State University. (2017). Soldier Beetle. Western Colorado Insects. https://wci.extension.colostate.edu/wp-content/uploads/sites/14/2017/03/SoldierBeetles.pdf.</ref> | <ref name= "colorado">Colorado State University. (2017). Soldier Beetle. Western Colorado [[Insects]]. https://wci.extension.colostate.edu/wp-content/uploads/sites/14/2017/03/SoldierBeetles.pdf.</ref> | ||
<ref name= "bugguide">BugGuide. (2025). Family Cantharidae - Soldier Beetles. BugGuide. https://bugguide.net/node/view/118/tree.</ref> | <ref name= "bugguide">BugGuide. (2025). Family Cantharidae - Soldier Beetles. BugGuide. https://bugguide.net/node/view/118/tree.</ref> | ||
<ref name="ucipm">UCIPM. (2025). Soldier Beetles. UCIPM. https://ipm.ucanr.edu/natural-enemies/soldier-beetles/#gsc.tab=0.</ref> | <ref name="ucipm">UCIPM. (2025). Soldier Beetles. UCIPM. https://ipm.ucanr.edu/natural-enemies/soldier-beetles/#gsc.tab=0.</ref> | ||
Line 55: | Line 57: | ||
<ref name="catron">Catron, K.A.. (2021). Biology of Chauliognathus spp. (Hentz) (Coleoptera: Cantharidae) in Virginia agroecosystems. Virginia Polytechnic Institute and State University. https://vtechworks.lib.vt.edu/server/api/core/bitstreams/f6f5fd1b-f813-4947-a14a-1b06d989fc18/content.</ref> | <ref name="catron">Catron, K.A.. (2021). Biology of Chauliognathus spp. (Hentz) (Coleoptera: Cantharidae) in Virginia agroecosystems. Virginia Polytechnic Institute and State University. https://vtechworks.lib.vt.edu/server/api/core/bitstreams/f6f5fd1b-f813-4947-a14a-1b06d989fc18/content.</ref> | ||
<ref name="podabrus">Montana State University. (2022). Podabrus sp. Montana State University. https://www.montana.edu/yellowstoneinsects/coleoptera/cantharidae/podabrus_sp.html#:~:text=Podabrus%20is%20a%20genus%20in,exclusively%20near%20bodies%20of%20water.</ref> | <ref name="podabrus">Montana State University. (2022). Podabrus sp. Montana State University. https://www.montana.edu/yellowstoneinsects/coleoptera/cantharidae/podabrus_sp.html#:~:text=Podabrus%20is%20a%20genus%20in,exclusively%20near%20bodies%20of%20water.</ref> | ||
<ref name="pelletier">Pelletier G. & Hebert, C. (2014). The Cantharidae of Eastern Canada and Northeastern United States. Canadian Journal of [[Arthropod]] Identification. https://doi.org/10.3752/cjai.2014.25.</ref> | |||
<ref name="crowley">Crowley, L., Chua, P., & Kusy, D.. (2023). The genome sequence of a soldier beetle, Podabrus alpinus (Paykull, 1798). PubMed Central. https://doi.org/10.12688/wellcomeopenres.18890.1.</ref> | |||
<ref name="minnesota">MinnesotaSeasons.com. (2025). Wrinkled Soldier Beetle. MinnesotaSeasons.com. http://www.minnesotaseasons.com/Insects/wrinkled_soldier_beetle.html#:~:text=Podabrus%20rugosulus%20is%20a%20small,and%20rounded%20at%20the%20tip.</ref> | |||
<ref name= "wildlife">The Wildlife Trusts. (2025). Soldier beetle. The Wildlife Trusts. https://www.wildlifetrusts.org/wildlife-explorer/invertebrates/beetles/soldier-beetle.</ref> | |||
<ref name= "pics">iNaturalist. https://www.inaturalist.org/.</ref> | <ref name= "pics">iNaturalist. https://www.inaturalist.org/.</ref> |
Latest revision as of 15:18, 2 May 2025
Description & Overview
![]() | ||||||
---|---|---|---|---|---|---|
Kingdom | Phylum | Subphylum | Class | Order | Suborder | Family |
Animalia | Arthropoda | Hexapoda | Insecta | Coleoptera | Polyphaga | Cantharidae |
Soldier beetles (Cantharidae) are the second largest group in the beetle (Coleoptera) superfamily Elateroidea, containing approximatley 5,500 species. These beetles are common throughout the temperate region of the northern hemisphere [3].
Soldier beetle larvae tend to be darkly colored and look almost reptilian. As adults, soldier beetles grow to about half of an inch long and are yellow, orange, brown, or black in color [4].
Though soldier beetles are sometimes considered pests, they are beneficial, as they are a pollinator species [4]. Soldier beetles may be confused with certain species of blister beetles, fireflies, or glowworm beetles [5].
Ecology & Evolution
Soldier beetles have four life stages: Egg, Larva, Pupa, and Adult. Eggs are laid from late August to early September. Larvae are assumed to predate on small invertebrates within the soil. After winter, larvae are found moving through leaf litter, plant debris, and other areas of high humidity. Pupae form in early summer, and adults emerge starting in late July [4]. One notable adaption of soldier beetles is that select ones can produce compounds to help stave off predators, like spiders [6].
From midsummer to early fall, soldier beetles tend to pollinate yellow flowers of the family Asteraceae (such as dandelions and goldenrods). Some adult soldier beetles feed on nectar and pollen [6]. Other adults may prey on aphids and soft-bodied insects [5].
Along with their important ecological role, soldier beetles have been significant in evolutionary studies. Currently, 25 species in 16 genera of soldier beetles have been discovered in fossiliferous amber resin. The oldest soldier beetle fossil discovered comes from Early Cretaceous Lebanese amber [7].
Based on molecular dating, the Cantharidae family is thought to have originated 156.3 million years ago [3].
Common Soldier Beetle Genera
Cantharis

Cantharis species are usually grayish-brown with a red or orange head and prothorax [6]. Beetles in this genus are commonly found in grasslands and forb fields [8]. Cantharis species are some of the largest soldier beetles, and adults can be between 9 to 13 millimeters long. These beetles have a red pronotum (back of head) with a black marking in the center. The top of these beetles' heads, as well as their wing covers, are black [9].
Chauliognathus

Most Chauliognathus species are found in the western United States, Australia, and South America, excepting two species in the state of Virginia. Adults in this genus are also some of the largest soldier beetles at 8 to 15 millimeters long. They are yellow to orange, with black coloration on their forewings and prothorax [10]. Like Cantharis, Chauliognathus species are also commonly found in grasslands and forb fields [8].
Podabrus

Species in the Podabrus genus tend to feed on aphids and soft-bodied insects [11]. The larve of Podabrus genus are thought to reside in organic debris and the upper layers of soil [12]. Podabrus adults typically range from 7 to 9 millimeters long. They tend to have leathery, flexible wing covers and are usually brownish-black with yellow to red sides and yellow faces [13].
References
[3] [4] [6] [1] [5] [7] [10] [11] [8] [12] [13] [9] [2]
- ↑ Jump up to: 1.0 1.1 BugGuide. (2025). Family Cantharidae - Soldier Beetles. BugGuide. https://bugguide.net/node/view/118/tree.
- ↑ Jump up to: 2.0 2.1 2.2 2.3 2.4 iNaturalist. https://www.inaturalist.org/.
- ↑ Jump up to: 3.0 3.1 3.2 Motyka, M., Kusy, D., Biffi, G., Geiser, M., Kazantsev, S.V., Bilkova, R., Jahodarova, E., Vogler, A.P., Bocak, L.. (2023). Untangling the evolution of soldier beetles (Coleoptera: Cantharidae) and the evaluation of the morphological phylogenetic signal in a soft-bodied elateroid lineage. Cladistics. 39: 548-570. https://doi.org/10.1111/cla.12555.
- ↑ Jump up to: 4.0 4.1 4.2 4.3 Hahn, Jeffrey. (2023). Soldier beetles. University of Minnesota Extension. https://extension.umn.edu/yard-and-garden-insects/soldier-beetles#:~:text=Soldier%20beetles%20are%20beneficial%20insects&text=Because%20of%20their%20frequent%20contact,go%20away%20on%20their%20own.
- ↑ Jump up to: 5.0 5.1 5.2 UCIPM. (2025). Soldier Beetles. UCIPM. https://ipm.ucanr.edu/natural-enemies/soldier-beetles/#gsc.tab=0.
- ↑ Jump up to: 6.0 6.1 6.2 6.3 Colorado State University. (2017). Soldier Beetle. Western Colorado Insects. https://wci.extension.colostate.edu/wp-content/uploads/sites/14/2017/03/SoldierBeetles.pdf.
- ↑ Jump up to: 7.0 7.1 Entomology Today. New Soldier Beetle Species Found in 99-million-year-old Amber. Entomological Society of America. https://entomologytoday.org/2016/10/14/new-soldier-beetle-species-found-in-99-million-year-old-amber/.
- ↑ Jump up to: 8.0 8.1 8.2 Pelletier G. & Hebert, C. (2014). The Cantharidae of Eastern Canada and Northeastern United States. Canadian Journal of Arthropod Identification. https://doi.org/10.3752/cjai.2014.25.
- ↑ Jump up to: 9.0 9.1 The Wildlife Trusts. (2025). Soldier beetle. The Wildlife Trusts. https://www.wildlifetrusts.org/wildlife-explorer/invertebrates/beetles/soldier-beetle.
- ↑ Jump up to: 10.0 10.1 Catron, K.A.. (2021). Biology of Chauliognathus spp. (Hentz) (Coleoptera: Cantharidae) in Virginia agroecosystems. Virginia Polytechnic Institute and State University. https://vtechworks.lib.vt.edu/server/api/core/bitstreams/f6f5fd1b-f813-4947-a14a-1b06d989fc18/content.
- ↑ Jump up to: 11.0 11.1 Montana State University. (2022). Podabrus sp. Montana State University. https://www.montana.edu/yellowstoneinsects/coleoptera/cantharidae/podabrus_sp.html#:~:text=Podabrus%20is%20a%20genus%20in,exclusively%20near%20bodies%20of%20water.
- ↑ Jump up to: 12.0 12.1 Crowley, L., Chua, P., & Kusy, D.. (2023). The genome sequence of a soldier beetle, Podabrus alpinus (Paykull, 1798). PubMed Central. https://doi.org/10.12688/wellcomeopenres.18890.1.
- ↑ Jump up to: 13.0 13.1 MinnesotaSeasons.com. (2025). Wrinkled Soldier Beetle. MinnesotaSeasons.com. http://www.minnesotaseasons.com/Insects/wrinkled_soldier_beetle.html#:~:text=Podabrus%20rugosulus%20is%20a%20small,and%20rounded%20at%20the%20tip.